Read the passage and answer the question. Passage: "Despite warnings from environmentalists, the coastal resort was built on a stretch of beach that served as a nesting ground for sea turtles. Within two years of opening, the number of turtle nests on that beach dropped by 80%. Environmentalists argued that artificial lighting confused the turtles during nesting season. The resort management installed red lights instead of white ones and reduced lighting after 10pm. The following year, turtle nesting numbers partially recovered." Question: In the sentence "Environmentalists argued that artificial lighting confused the turtles during nesting season", what does "the turtles" refer to?
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Correct answer: The sea turtles that nested on the beach where the resort was built
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